Agaricales

Inocybaceae

Inocybe

Inocybe abjecta P. Karst. (Kuyper, 1986; Bon, 1997)
= Inocybe caucasica Singer (Kuyper, 1986)

Inocybe acuta Boud.
= Inocybe acutella Bon
= Inocybe striata Bres.

Inocybe aeruginascens Babos

Inocybe albovelutipes Stangl

Inocybe alnea Stangl

Inocybe alpigenes (E. Horak) Bon

Inocybe amblyospora Kühner (Bandini et al., 2023)

Inocybe amethystina Kuyper (Bandini et al., 2021)

Inocybe appendiculata Kühner

Inocybe argenteolutea Vauras

Inocybe assimilata Britzelm.
= Astrosporina umbrina (Bres.) Rea
= Inocybe umbrina Bres.

Inocybe asterospora Quél.
≡ Clypeus asterosporus (Quél.) P. Karst.

Inocybe aurea Huijsman

Inocybe auricoma (Batsch) Sacc.

Inocybe borealis M. Lange

Inocybe bresadolae Massee

Inocybe brunnea Quél.

Inocybe brunneoatra (R. Heim) P.D. Orton
≡ Inocybe descissa var. brunneoatra R. Heim

Inocybe brunneotomentosa Huijsman

Inocybe caesariata (Fr.) P. Karst.

Inocybe calida Velen.
= Inocybe brunneorufa Stangl & J. Veselský

Inocybe calospora Quél.

Inocybe carelica Singer

Inocybe castanea Peck

Inocybe catalaunica Singer (Bandini, Oertel, et al., 2022)
= Inocybe abietis Kühner (Bandini, Oertel, et al., 2022)
= Inocybe leiocephala D.E. Stuntz

Inocybe cincinnata (Fr.) Quél. (Bandini et al., 2021)
≡ Inocybe cincinnata var. cincinnata (Fr.) Quél.
= Inocybe cristata (Scop.) J. Schröt.
= Inocybe squarrosoamethystina Singer (Kuyper, 1986)
Inocybe phaeocomis (Pers.) Kuyper sensu Kuyper 1986 (Bon, 1997; Bandini et al., 2021)
Inocybe phaeocomis var. phaeocomis (Pers.) Kuyper sensu Kuyper 1986 (Bon, 1997; Bandini et al., 2021)

Inocybe concinnula J. Favre

Inocybe confusa P. Karst.

Inocybe corydalina Quél.
= Inocybe corydalina var. corydalina Quél.

Inocybe crocifolia Herink (Bandini et al., 2021)
≡ Inocybe flocculosa var. crocifolia (Herink) Kuyper

Inocybe cryptocystis D.E. Stuntz

Inocybe curvipes P. Karst.
= Inocybe boltonii R. Heim
= Inocybe lanuginella (J. Schröt.) Konrad & Maubl.

Inocybe cylindrocystis G.F. Atk.

Inocybe decemgibbosa (Kühner) Vauras

Inocybe decipiens Bres. (Bandini, Brandrud, et al., 2022)

Inocybe dunensis P.D. Orton (Bandini, Brandrud, et al., 2022)

Inocybe egenula J. Favre

Inocybe eutheles (Fr.) Quél.

Inocybe euviolacea E. Ludw.
≡ Inocybe geophylla var. violacea (Pat.) Sacc.

Inocybe excoriata Peck

Inocybe favrei Bon

Inocybe fibrosa (Sowerby) Gillet

Inocybe fibrosoides Kühner

Inocybe flocculosa Sacc.
≡ Inocybe flocculosa var. flocculosa Sacc.
= Inocybe gausapata Kühner

Inocybe fraudans (Britzelm.) Sacc.

Inocybe fuligineoatra Huijsman

Inocybe fulvella Bres.

Inocybe furfurea Kühner

Inocybe geophylla (Bull.) P. Kumm. (Bandini et al., 2021)
≡ Inocybe geophylla f. geophylla (Bull.) P. Kumm.
≡ Inocybe geophylla var. geophylla (Bull.) P. Kumm.
≡ Agaricus geophyllus Bull.
= Agaricus geophyllus var. candidus Fr. (Kuyper, 1986)
= Inocybe xantholeuca Kuyper (Bandini et al., 2021)

Inocybe glabrescens Velen. (Bandini et al., 2021)

Inocybe glabripes Ricken (Bandini et al., 2021)
= Inocybe fuscidula Velen. (Bandini et al., 2021)
= Inocybe fuscidula var. fuscidula Velen. (Bandini et al., 2021)
= Inocybe microspora J.E. Lange (Bandini et al., 2021)
= Inocybe descissa var. microspora (J.E. Lange) R. Heim (Bandini et al., 2021)

Inocybe glabrodisca P.D. Orton

Inocybe godeyi Gillet

Inocybe goniopusio Stangl

Inocybe grammata Quél.

Inocybe griseolilacina J.E. Lange (Bandini et al., 2021)

Inocybe griseoscabrosa (Peck) Earle

Inocybe griseovelata Kühner (Bandini et al., 2021; Bandini, Brandrud, et al., 2022)

Inocybe helobia (Kuyper) Bandini, B. Oertel & U. Eberh.
≡ Inocybe lacera var. helobia Kuyper

Inocybe hirtella Bres. (Bandini, Oertel, et al., 2022)
≡ Inocybe hirtella var. hirtella Bres.
= Inocybe hirtella var. bispora Kuyper (Bandini, Oertel, et al., 2022)

Inocybe hypophaea Furrer-Ziogas (Bandini et al., 2021)

Inocybe hypotheja Kühner

Inocybe hystrix (Fr.) P. Karst.

Inocybe impexa (Lasch) Kuyper
= Inocybe maritima P. Karst.
= Agaricus maritimus Fr. 

Inocybe incarnata Bres.

Inocybe inodora Velen.

Inocybe intricata Peck
= Inocybe intricata var. pallidistipitata Grund & D.E. Stuntz

Inocybe lacera (Fr.) P. Kumm. (Bandini, Brandrud, et al., 2022)
≡ Inocybe lacera var. lacera (Fr.) P. Kumm.
≡ Agaricus lacerus Fr. 
= Inocybe lacera var. regularis Kuyper
= Inocybe lacera var. rhacodes (J. Favre) Kuyper

Inocybe langei R. Heim (Bandini, Oertel, et al., 2022)
= Inocybe langei f. major J.E. Lange
= Inocybe langei var. major Singer

Inocybe lanuginosa (Bull.) Kalchbr.
≡ Inocybe lanuginosa var. lanuginosa (Bull.) Kalchbr.
≡ Agaricus lanuginosus Bull.

Inocybe leptocystis G.F. Atk.

Inocybe leptophylla G.F. Atk.
= Inocybe casimirii Velen.

Inocybe lilacina (Peck) Kauffman
≡ Inocybe geophylla var. lilacina (Peck) Gillet
= Agaricus geophyllus Sowerby (Kuyper, 1986)

Inocybe luteipes J. Favre

Inocybe lutescens Velen. (Bandini et al., 2021)

Inocybe maculipes J. Favre

Inocybe margaritispora (Berk.) Sacc.
= Inocybe phaeosticta Furrer-Ziogas

Inocybe melanopoda D.E. Stuntz

Inocybe minuta P. Karst.

Inocybe mixtilis (Britzelm.) Sacc.

Inocybe moelleri Eyssart. & A. Delannoy

Inocybe monochroa J. Favre

Inocybe muricellata Bres.

Inocybe napipes J.E. Lange

Inocybe nematoloma Joss.

Inocybe nitidiuscula (Britzelm.) Lapl.
= Inocybe friesii R. Heim

Inocybe oblectabilis (Britzelm.) Sacc.

Inocybe obscurobadia (J. Favre) Grund & D.E. Stuntz (Bandini et al., 2021)

Inocybe obscuroides P.D. Orton
Inocybe phaeocomis var. major (S. Petersen) Kuyper sensu Kuyper 1986 (Bandini et al., 2021)

Inocybe ochracea Stangl

Inocybe ochroalba Bruyl. (Bandini, Oertel, et al., 2022)

Inocybe olida Maire

Inocybe oreina J. Favre

Inocybe ovatocystis Boursier & Kühner
≡ Inocybe lanuginosa var. ovatocystis (Boursier & Kühner) Stangl

Inocybe paludinella (Peck) Sacc.

Inocybe parvicystis Rodr.-Campo & Esteve-Rav.

Inocybe pelargonium Kühner

Inocybe petiginosa (Fr.) Gillet
≡ Hebeloma petiginosum (Fr.) P. Kumm.

Inocybe phaeodisca Kühner (Bandini et al., 2021)

Inocybe phaeoleuca Kühner
≡ Inocybe splendens var. phaeoleuca (Kühner) Kuyper

Inocybe posterula (Britzelm.) Sacc.

Inocybe praetervisa Quél.
≡ Astrosporina praetervisa (Quél.) J. Schröt.

Inocybe proximella P. Karst.

Inocybe pseudoasterospora Kühner & Boursier
= Inocybe pseudoasterospora var. microsperma Kuyper & P.-J. Keizer
= Inocybe pseudoasterospora var. microsperma Weholt

Inocybe pseudodestricta Stangl & J. Veselský

Inocybe pseudohiulca Kühner

Inocybe pseudoreducta Stangl & Glowinski

Inocybe pseudoumbrina Stangl

Inocybe pusio P. Karst. (Bandini et al., 2021)

Inocybe putilla Bres.

Inocybe pyriodora (Pers.) P. Kumm.
≡ Agaricus pyriodorus Pers.

Inocybe queletii Konrad

Inocybe radiata Peck

Inocybe ravaensis Kalamees & Shchukin

Inocybe relicina (Fr.) Quél.

Inocybe rennyi (Berk. & Broome) Sacc.

Inocybe rufoalba Sacc.
= Inocybe jacobi Kühner

Inocybe rufuloides Bon (Bandini et al., 2023)

Inocybe rupestris J. Favre

Inocybe salicis-herbaceae Kühner

Inocybe sambucina (Fr.) Quél.
≡ Agaricus sambucinus Fr. 

Inocybe sapinea Velen.

Inocybe serotina Peck

Inocybe similis Bres.

Inocybe sindonia (Fr.) P. Karst. (Bandini et al., 2021; Bandini, Oertel, et al., 2022)
= Inocybe kuehneri Stangl & J. Veselský (Bandini, Oertel, et al., 2022)

Inocybe soluta Velen. (Bandini, Brandrud, et al., 2022)
= Inocybe albofibrillosa Stangl & Schmid-Heckel (Bandini, Brandrud, et al., 2022)

Inocybe splendens R. Heim
= Inocybe phaeoleuca var. phaeoleuca Kühner
= Inocybe splendens var. splendens R. Heim

Inocybe squarrosa Rea

Inocybe stellatospora (Peck) Massee

Inocybe subargentea Singer

Inocybe subcarpta Kühner & Boursier (Bandini, Brandrud, et al., 2022)
= Inocybe brevispora Huijsman (Bandini, Brandrud, et al., 2022)

Inocybe subnudipes Kühner (Bandini, Brandrud, et al., 2022)

Inocybe subochracea (Peck) Earle

Inocybe subpaleacea Kühner

Inocybe tabacina Furrer-Ziogas

Inocybe tarda Kühner (Bandini et al., 2021)

Inocybe taxocystis (J. Favre & E. Horak) Senn-Irlet

Inocybe tenebrosa Quél.
= Inocybe atripes G.F. Atk.

Inocybe teraturgus M.M. Moser

Inocybe terrifera Kühner

Inocybe tigrina R. Heim (Bandini et al., 2021)

Inocybe tjallingiorum Kuyper (Bandini et al., 2021)

Inocybe umbratica Quél.

Inocybe urceolicystis Stangl & Vauras

Inocybe ventricosa G.F. Atk.

Inocybe virgatula Kühner (Bandini et al., 2021)

Inocybe vulpinella Bruyl.

Inocybe whitei (Berk. & Broome) Sacc.
= Inocybe geophylla var. lateritia (Berk. & Broome) W.G. Sm.
= Inocybe pudica Kühner

Inocybe xanthomelas Boursier & Kühner

Inosperma

Inosperma adaequatum (Britzelm.) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe adaequata (Britzelm.) Sacc.
= Inocybe jurana (Pat.) Sacc. (Kuyper, 1986)

Inosperma bongardii (Weinm.) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Agaricus bongardii Weinm.
≡ Inocybe bongardii (Weinm.) Quél.
≡ Inocybe bongardii var. bongardii (Weinm.) Quél.
= Agaricus gratus Weinm. (Kuyper, 1986)

Inosperma calamistratum (Fr.) Matheny & Esteve-Rav. (Matheny et al., 2020; Esteve-Raventós et al., 2024)
≡ Inocybe calamistrata (Fr.) Gillet
≡ Agaricus calamistratus Fr. 
= Inocybe hirsuta (Lasch) Quél. (Kuyper, 1986; Esteve-Raventós et al., 2024)
= Inosperma hirsutum (Lasch) Matheny & Esteve-Rav. (Kuyper, 1986; Esteve-Raventós et al., 2024)

Inosperma cervicolor (Pers.) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe cervicolor (Pers.) Quél.
= Inocybe corrubescens Singer (Kuyper, 1986)

Inosperma cookei (Bres.) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe cookei Bres.
= Inocybe cookei var. cookei Bres. (Matheny et al., 2020)

Inosperma erubescens (A. Blytt) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe erubescens A. Blytt
= Inocybe patouillardii Bres. (Kuyper, 1986)

Inosperma fastigiellum (G.F. Atk.) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe fastigiella G.F. Atk.

Inosperma geraniodorum (J. Favre) Matheny & Esteve-Rav. (Matheny et al., 2020; Esteve-Raventós et al., 2024)
≡ Inocybe geraniodora J. Favre

Inosperma kuthanii (Stangl & J. Veselský) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe cookei var. kuthanii (Stangl & J. Veselský) Kuyper

Inosperma maculatum (Boud.) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe maculata Boud.

Inosperma praetermissum (P. Karst.) Esteve-Rav., E. Larss. & Pancorbo (Esteve-Raventós et al., 2024)
≡ Inocybe praetermissa P. Karst.

Inosperma quietiodor (Bon) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe quietiodor Bon

Inosperma reisneri (Velen.) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe reisneri Velen.

Inosperma rimosoides (Peck) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe rimosoides Peck

Inosperma subhirsutum (Kühner) Matheny & Esteve-Rav. (Matheny et al., 2020; Esteve-Raventós et al., 2024)
≡ Inocybe subhirsuta Kühner

Mallocybe

Mallocybe agardhii (N. Lund) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe agardhii (N. Lund) P.D. Orton
≡ Pholiota agardhii (N. Lund) M.M. Moser

Mallocybe arthrocystis (Kühner) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe arthrocystis Kühner

Mallocybe dulcamara (Pers.) Vizzini, Maggiora, Tolaini & Ercole
≡ Inocybe dulcamara (Pers.) P. Kumm.
≡ Inocybe dulcamara f. dulcamara (Pers.) P. Kumm.
≡ Inocybe dulcamara var. dulcamara (Pers.) P. Kumm.
= Inocybe dulcamara f. solida J. Favre

Mallocybe fulvipes (Kühner) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe fulvipes Kühner

Mallocybe fuscomarginata (Kühner) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe fuscomarginata Kühner

Mallocybe gymnocarpa (Kühner) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe gymnocarpa Kühner

Mallocybe heimii (Bon) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe heimii Bon

Mallocybe leucoblema (Kühner) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe leucoblema Kühner

Mallocybe malenconii (R. Heim) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe malenconii R. Heim

Mallocybe perbrevis (Weinm.) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Agaricus perbrevis Weinm.
≡ Inocybe perbrevis (Weinm.) Gillet

Mallocybe peronata (J. Favre) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe dulcamara f. peronata J. Favre

Mallocybe pygmaea (J. Favre) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe pygmaea (J. Favre) E. Ludw.
≡ Inocybe dulcamara f. pygmaea J. Favre

Mallocybe squamosoannulata (J. Favre) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe squamosoannulata (J. Favre) E. Ludw.
≡ Inocybe dulcamara var. squamosoannulata (J. Favre) Schmid-Heckel

Mallocybe substraminipes (Kühner) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe substraminipes Kühner

Mallocybe terrigena (Fr.) Matheny, Vizzini & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe terrigena (Fr.) Kuyper
≡ Inocybe terrigena (Fr.) Kühner

Mallocybe umbrinofusca (Kühner) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe umbrinofusca Kühner

Pseudosperma

Pseudosperma curreyi (Berk.) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe curreyi (Berk.) Sacc.
≡ Inocybe fastigiata subsp. curreyi (Berk.) Dermek & J. Veselský

Pseudosperma flavellum (P. Karst.) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe flavella P. Karst.
≡ Inocybe flavella var. flavella P. Karst.
= Inocybe flavella var. roseipes (Bon) Kuyper

Pseudosperma obsoletum (Quadr.) Valade (Matheny et al., 2020)
≡ Inocybe obsoleta (Quadr.) Valade
≡ Inocybe obsoleta Romagn.

Pseudosperma perlatum (Cooke) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe perlata (Cooke) Sacc.

Pseudosperma rimosum (Bull.) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe rimosa (Bull.) Kalchbr.
≡ Agaricus rimosus Bull.
= Inocybe fastigiata f. alpestris R. Heim
= Inocybe destricta (Fr.) Quél.
= Inocybe fastigiata (Schaeff.) Kalchbr.
= Inocybe fastigiata f. fastigiata (Schaeff.) Kalchbr.
= Inocybe fastigiata f. subcandida Malençon

Pseudosperma sororium (Kauffman) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe sororia Kauffman

Pseudosperma squamatum (J.E. Lange) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe squamata J.E. Lange

Pseudosperma umbrinellum (Bres.) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe umbrinella Bres.

Pseudosperma vinosistipitatum (Grund & D.E. Stuntz) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe vinosistipitata Grund & D.E. Stuntz

Pseudosperma xanthocephalum (P.D. Orton) Matheny & Esteve-Rav. (Matheny et al., 2020)
≡ Inocybe xanthocephala P.D. Orton

References

Bandini D., Bandini G., Bensch K., Campo F. J. R., Oertel B., Valade F., Eberhardt U. 2023. Three new smooth-spored species of Inocybe, two new epitypes, and updates on earlier epitypifications // Integrative Systematics: Stuttgart Contributions to Natural History 6(2): 1–37. https://doi.org/10.18476/2023.787646

Bandini D., Brandrud T. E., Dima B., Dondl M., Fachada V., Hussong A., Mifsud S., Oertel B., Campo F. J. R., Thüs H., Vauras J., Weholt Ø., Eberhardt U. 2022. Fibre caps across Europe: Type studies and 11 new species of Inocybe (Agaricales, Basidiomycota) // Integrative Systematics: Stuttgart Contributions to Natural History 5(2): 1–85. https://doi.org/10.18476/2022.901982

Bandini D., Oertel B., Eberhardt U. 2021. A fresh outlook on the smooth-spored species of Inocybe: Type studies and 18 new species // Mycological Progress 20(9): 1019–1114. https://doi.org/10.1007/s11557-021-01712-w

Bandini D., Oertel B., Eberhardt U. 2022. More smooth-spored species of Inocybe (Agaricales, Basidiomycota): Type studies and 12 new species from Europe // Persoonia - Molecular Phylogeny and Evolution of Fungi 48(1): 91–149. https://doi.org/10.3767/persoonia.2022.48.03

Bon M. 1997. Clé monographique du genre Inocybe (Fr.) Fr. (2ème partie: sous-genre Inocybe = Inocybium (Earle) Sing.) // Documents Mycologiques 27(108): 1–77.

Esteve-Raventós F., Larsson E., Pancorbo F., Bizio E., Altés A., Turégano Y., Moreno G., Olariaga I. 2024. A Taxonomic and Phylogenetic Contribution on Inosperma Section Inosperma (Agaricales, Inocybaceae) in Europe: Calamistratum and Geraniodorum Groups // Journal of Fungi 10(6): 374. https://doi.org/10.3390/jof10060374

Kuyper Th. W. 1986. A revision of the genus Inocybe in Europe. I. Subgenus Inosperma and the smooth-spored species of subgenus Inocybe // Persoonia - Supplement 3: 1–247.

Matheny P. B., Hobbs A. M., Esteve-Raventós F. 2020. Genera of Inocybaceae: New skin for the old ceremony // Mycologia 112(1): 83–120. https://doi.org/10.1080/00275514.2019.1668906